I got this interpretation when I watched the PV with the lyrics. They were very in-sync with the PV!
The song talks about a girl who's fallen head over heels about this guy, who's crush quickly develops into an obsession where then she asks herself (or the audience) whether or not they will try their hardest to be themselves or simply fall into the trap of this one sided "love" (this part is sung in the bridge).
In short, the four Ayu's represent the different progressive stages of affection and response that the girl goes through as in the lyrics portray.
blue ayu: (initial) flirty, innocence, just playing around
Quote:
Talking about nothing at all,
We acted in a moment and displayed our habits.
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black ayu: (second verse) the darker, real ayu symbolizing her progression of obsession
Quote:
"Where are you, what are you doing around this time?"
Right after I thought such things,
Even for just a moment,
I wanted you to remember me.
I wished for YOU and ME
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white ayu: (second chorus) confusion over whether to put her head before her heart or vice versa
red ayu: (bridge) portrays the girl/image of the girl who gave into her obsession over the boy, who asks Ayu if she'll be strong or give in aswell.
Quote:
The gap between idealism and reality
Is so very cutting
And as we cannot cut them off, they are opening up.
In this place, will we give up?
Or will we persevere and do our very best?
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The chorus reinforces these ideas of being obsessive and out of mind which are complemented with the 1.2.3.4.'s creating this child-like effect.
Quote:
SO 1.2.3.4, for a while
1.2 STEP you, sometime soon
I am becoming greedy
1.2.3.4, for a while
1.2 STEP you, sometime soon
Will I begin to want only your smile?
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Whilst the robot man at the beginning might represent the man that she wants (the rotating thingo showcasing the different ayu's might be the girl saying "which of me do you like?") and finally the resolution of the girl's inner conflict is visible with the climax of the PV with the rain falling onto the "complete" ayu (rain often symbolizes salvation) and the other four ayu disappears - symbolizing that she got over him.
Anyhow that's what I got from watching the PV with the translated lyrics on - you should rewatch it to with subtitles to get a clearer idea!